I have replied to you in the support thread you have opened, you can check my response there. For your convenience I'm pasting my reply below:
" Can you, please, inform me about the exact scenario where this problem reproduces? Generally there might be a reason for default permissions to differ between upgraded version and a blank project created with 4.1 Project Manager - we did introduce some changes in the permissions for the default roles in 4.1 but the upgrade script should not change the permissions on an already running site, that's why it should be disabled during upgrade. Can you, please verify if the issue reproduces with a blank project after upgrade, and provide me with some more details so that I can inspect this issue further. Thanks in advance."
Looking forward for your feedback on this, if you have any additional questions, please do not hesitate to write back.
All the best,
the Telerik team